In pictures: Galapagos tortoises head home after decades in captivity

Galapagos National Park rangers release chelonidis hoodensis turtles in the area called Las Tunas, 2.5 km from the coast of Espanola Island in the Galapagos archipelago, Ecuador. A group of giant tortoises who have lived in captivity for decades and helped rescue their species from the brink of extinction were released into the wild in the Galapagos Islands.

Giant tortoises stand moments after release to their original habitat on Isla Espanola. The tortoises were bred in captivity to repopulate the islands after their numbers in the wild went down to just 15 individuals, said park director Danny Rueda. There are now more than 2,000 of them on Espanola Island, he said.

Park rangers move chelonidis hoodensis turtles before being released. The giant tortoises are known for their long, leathery necks and lifespans of over 100 years. Virtually synonymous with the Galapagos Islands off the coast of Ecuador, they were one of the species that helped Charles Darwin form his theory of evolution in the 19th century.

One of the newly freed tortoises is Diego (pictured), estimated to be around 100 years old. Renowned for his fertility, he alone has fathered some 800 offspring.

Diego's return to his native Espanola comes after around eight decades of living in a California zoo and then in the tortoise recovery program on Santa Cruz island in the Galapagos.

For his final journey home, Diego was taken by boat to Espanola, along with 14 of his companions. From there, rangers used backpacks to take the tortoises - who can weigh up to 180 kg - to an area where cactus grows in abundance that will help them readapt.

There, they will be monitored with GPS trackers. It has been a conservation success story.

"We can shut down the captive breeding program of this species because their natural behaviour is effective," said Rueda.

Volunteer from the Galapagos National Park carrying giant tortoises to be released on the Isla Espanola.

Ranger Carlos Gaona carries a tortoise named Diego, to return him to his natural habitat on Isla Española, Galapagos National Park.